The Howling Commandos were an elite combat unit that had been led by Captain America during World War II. At the end of the war and Captain America's apparent demise, their unit continued to be active and was led by Dum Dum Dugan.
The new unit, led by Rogers and Bucky Barnes, traveled around Western Europe to eliminate every HYDRA base they could find. Their operations were some of the most successful of the entire war. The Howling Commandos were responsible for the capture of Arnim Zola during a raid on a train in the Alps. It was on this mission that Bucky Barnes was lost and presumed dead.
Following World War II the Commandos served as a tactical team for the 107th Infantry Regiment. Some members eventually leaving the unit but new recruits came. In 1946, the Howling Commandos were called by Peggy Carter to help her, SSR agent Jack Thompson, and his team in infiltrating a Leviathan base camp in Byelorussia. They met Carter and the team and escorted them to Belarus. Infiltrating the facility, they discovered various methods of dispensing subliminal messages. They encountered a little girl supposedly left behind, and Dum Dum Dugan went to comfort her, only to be non-fatally stabbed by the girl, who quickly shot and killed Junior Juniper before escaping.
Going deeper into the facility, the team found the scientists Nikola and Ivchenko. Freeing them, they ran into an ambush where Nikola tried to use Happy Sam Sawyer as a hostage. Ivchenko killed Nikola, allowing the Commandos and Carter's team to escape.
The Howling Commandos said goodbye to Carter and her team, with Dugan giving Ivchenko his bottle of bourbon.
"Barnes is the only Howling Commando to give his life in service of his country."