The Chitauri Command Center was the large command ship that controlled all the Chitauri participating in the Chitauri Invasion of Earth. The command center was destroyed by Iron Man, ending the Battle of New York.
The Chitauri rode and held their army on the ship as they prepared for the Battle of New York. The battle ended when Iron Man took a nuclear missile that was meant to hit Manhattan and redirected it through the portal linking Earth to Thanos' domain. The nuclear missile then hit the ship, which controlled every Chitauri and Leviathan, causing them to die all at once.