Hive

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"Thousands of years ago, an Inhuman was born on this planet that was destined to rule it, so powerful, so fearsome that others were consumed with dread, and so they banished it from the Earth, sent it through the portal to a distant planet. HYDRA was founded with the sole purpose of engineering its return."

- Gideon Malick It is a parasitic Inhuman capable of infesting and possessing the bodies of the dead. Because of its great powers, It was banished to a barren planet. It's worshipers founded a society with the sole purpose of bringing It back to Earth to conquer it. Over the centuries, that society evolved into HYDRA. Eventually, It possessed Grant Ward's dead body when he was killed by Phil Coulson with his prosthetic hand on Maveth and was free on Earth once more.

Banished
Thousands of years ago, It was an Inhuman that was transformed by Terrigenesis into a parasitic organism of incredible power, capable of infesting the bodies of the dead and reanimating them into host bodies. Soon, other Inhumans grew so fearful of It that they banished It to another world through the use of a portal. It's followers, believing it to be destined to rule the world, soon organized a society with the purpose to bring their leader back to Earth. Over the centuries the society evolved into HYDRA.

The world upon which It was banished, Maveth, was once home to an extraterrestrial civilization, and was believed to have once been a beautiful paradise. However, a cataclysm that was theorized to have been caused by It destroyed the native species and reduced Maveth to a desolate wasteland.

Maveth War
"They had a chance to become something great, something beautiful, but in the end..." "How do you know so much about it?" "Because I was there."

- It and Leo Fitz

It was on Maveth at the time when a war was between the nine cities of the highly advanced civilization of Maveth. He saw how they fought each other and eventually destroyed their own race.

Centuries of Killing
"That THING has been killing everyone that's come through here for centuries." "How long have people known about the Monolith? And why keep sending people when there's no way back? It's as if they're being sacrificed!"

- Will Daniels and Jemma Simmons

For centuries, people were sent to the planet through the Monolith as ritualistic sacrifices, meeting their deaths at the hands of It and becoming vessels so that It could preserve itself within their bodies, sparing it from the decay of time. This was evidenced by the sword, music box and bag containing a bottle that Jemma Simmons found there in 2015, all of which were from bygone eras. Among these victims was Lord Manzini, who was sent when he grabbed the white stone in a lottery held by English lords and nobles in a castle in Gloucestershire, England in 1839. According to Lord Thornally, none who had entered through the Monolith by that point had ever returned.

Arrival of NASA
"It has this way of getting inside your head and just making you crazy. Austin threw himself off a cliff. Brubaker set himself on fire. Taylor took an axe to the gear and then he came after me."

- Will Daniels

By 2001, the Monolith was in the possession of NASA who sent a crew consisting of astronauts Will Daniels, Austin, Brubaker and Taylor to the planet. Once It became aware of their presence, It caused Austin, Brubaker and Taylor to lose their sanity. Austin and Brubaker both committed suicide, while Taylor attempted to kill Daniels, who killed him in self-defense. Daniels noted that they all entered a certain area before they went insane, where he discovered a mass grave and dubbed it the "No-Fly Zone". For the next 14 years, he survived by avoiding and occasionally outsmarting It, along with just being lucky.

Encounter with Jemma Simmons
"You were right, Will. I saw It, whatever it is. Something was out there."

- Jemma Simmons

In 2015, S.H.I.E.L.D. scientist Jemma Simmons was accidentally transported to Maveth while studying the Monolith. She took shelter with Daniels and did not believe his tales about It, crediting the frequent sandstorms to the planet's desert environment. She spent over two months trying to find a way to send them back to Earth, after which she went to search for answers in the "No-Fly Zone" despite Daniels' warnings. She was caught in a sandstorm, in which she saw It, its host body concealed by a dark cloak, approaching her. She ran back to the hovel in which she and Daniels had been hiding, realizing that what Daniels had told her about It was true.

Unruly Captives
"That's more like 100 meters. Unless there was some significant earthquake..." "It doesn't want us to leave."

- Jemma Simmons and Will Daniels

Simmons discovered that the stars were the key to discovering the location of a wormhole, and she came up with a plan to send a message to S.H.I.E.L.D. so that she and Daniels could be rescued. Once she had deduced the location of the wormhole and the time at which it would appear, they made a plan to place the message in a bottle and launch it through the wormhole with a grappling hook. However, It widened the canyon from 30 meters to 100 meters, making the task significantly more difficult. Daniels fired the bottle at the wormhole, missing it by a second. With the help of Elliot Randolph, Simmons' S.H.I.E.L.D. comrades Leo Fitz, Phil Coulson, Daisy Johnson, Alphonso Mackenzie and Bobbi Morse managed to activate the Monolith from the HYDRA Castle. Fitz fired a flare through it, which was spotted by Simmons and Daniels, who were awaiting the sunrise. In an attempt to prevent them from escaping, It summoned a sandstorm, separating them. It then appeared before Simmons in the body of an astronaut, fooling her into thinking that NASA had come to rescue Daniels. She alerted Daniels, who warned her that it was a trick. Daniels then told her to run while he stayed behind to protect her, using his last remaining bullet against It. Simmons was rescued by Fitz, and Daniels did not survive his encounter with It, suffering a large open wound in his leg during the battle. It then took possession of his corpse and accessed Daniels' memories to learn of Fitz and Simmons, also locating Daniels' hideout. It tossed Daniels' gun aside as it witnessed the planet's first sunrise in 18 years before It walked back to Daniels' hideout where It stayed and continued to await it's rescue.

Discovered by Ward's Team
"How long you been here?" "Feels like forever."

- Grant Ward and It

Following the death and possession of Will Daniels' corpse, It took refuge in Daniels' hideout. A month after Daniels' death, It was found by Leo Fitz, who It recognised from Daniels' memories. Fitz was followed by a HYDRA team led by Grant Ward. Though Ward wanted to kill "Daniels", Leo Fitz convinced Ward to spare him by explaining that Daniels knew the terrain of the planet and was vital to their journey. Walking ahead of the HYDRA team, It learned from Fitz that Ward wanted to bring the Inhuman back to Earth, and that they would both be kept alive in order to get to the extraction point. It then took the group through the "No-Fly Zone". As they traveled, Ward asked It about the ruins of a large HYDRA shrine the group saw in the distance when they first arrived on the planet, and It said that it used to live there. Ward explained to it that Earth had changed since he left, as the Inhumans had rapidly increased in numbers across the globe after a Terrigen Mist outbreak occurred. The conversation was cut short when It quietly caused a sudden sandstorm, which stopped their journey and forced Ward and his team to take over, with It refusing to accept Fitz's goggles. As the sandstorm raged on, the group was split up, hiding behind various large rock formations. It used this opportunity to stealthily kill two HYDRA agents, stabbing them in the back of the neck and leaving their bodies in the dirt, allowing It to flee with Fitz under the cover of the storm. By the time Ward and the remaining team stumbled upon the corpses of their colleagues, Fitz and It were gone.

It is Revealed
"You're not Will." "No. Will died saving Jemma ... from me."

- Leo Fitz and It

As Leo Fitz and It walked closer and closer to the portal's location, Fitz stopped when he noticed the ruins of the ancient civilization in the distance, baffled that Jemma Simmons never mentioned it. It told Fitz that Simmons never saw the ruins because it was in the "No-Fly Zone". It explained that Maveth was home to nine cities occupied by advanced beings that "feared change" and destroyed their own race through war. Fitz tried to bandage up It's injured leg, but was shocked to find most of the flesh had rotted away to the bone. As realization set in, It explained to Fitz that it was not really Will Daniels, as It killed Daniels during Simmons's escape from the planet. Fitz lunged for his weapon but was attacked by It. It was still able to match Fitz in a physical fight, overpowering him. Phil Coulson, who had followed HYDRA to Maveth, and the captive Grant Ward arrived at their location just as It prepared to bring down a large rock on Fitz' head. Coulson shot It twice before it could kill Fitz, and It collapsed to the ground. Despite being shot multiple times, It got up and staggered towards the now open portal. Fitz grabbed his gun and shot It multiple times, and once again It buckled to the ground. Moments later, however, It started to lift itself up, Fitz grabbed a flare gun and shot a flare into It's back, engulfing It in flames, and It collapsed as Will Daniels' body burned away. In the meantime, Coulson fought Ward and used his bionic hand to crush his ribcage. He then removed his bionic hand, threw it down next to Ward's corpse, and made his way to the portal with Fitz. It emerged from the burnt skull of Daniels and possessed Ward's corpse with Coulson's bionic hand in tow.

Meeting Gideon Malick
"Well I'll be damned."

- Gideon Malick

Despite S.H.I.E.L.D.'s attempt to destroy the remaining HYDRA forces and the last fragments of the Monolith with an airstrike attack against the HYDRA Castle, It managed to travel through the portal in time and made it out of the Castle before it was destroyed. Now on Earth, It, in possession of the body of Grant Ward, found and intercepted Gideon Malick on his way to the airport.

Seeing Modern-Day Earth
"They have advanced, but they haven't changed." "They?" "Humans. But you're not human anymore, are you? You don't believe I am what they say. Once I'm strong enough... You will."

- It and Giyera

Three months after his return to Earth, It was still resting in Gideon Malick's secret compound in Europe, trying to regain his strength, as its years trapped on Maveth had left it weak and slowly dying, and Grant Ward's body was rotting away. Eventually It built up the strength to speak and demanded to see Malick. When Malick entered he asked It was it needed, so the Inhuman demanded that he bring him food. Feeding on raw meat, It was watching numerous documentaries to learn about the technological advance of the mankind. Seeing the videos of several 20th Century wars, It told Giyera that humans haven't changed at all, they were still the same like in his time, if slightly more technologically advanced. When Giyera commented on It's recovery since the Battle on Maveth, It noted that Ward's body was dead and it had been dying. It then noted that Giyera himself was clearly an Inhuman, seemingly sensing Giyera's power from within before continuing to eat the meat and recover. Continuing to watch news reports, It listened as Malick and Giyera discussed the capture of Lucio, another Inhuman found in Bogotá. Hearing that Malick was overall disappointed by how his plan was unfolding, It turned to face them, commented on how Malick was losing his faith in It's power, with Malick noting that since It had failed to kill Phil Coulson, maybe his power was not as strong as he had hoped. It, however, vowed that Malick would believe; beginning to demonstrate his powers by turning Ward's skin to sand and sending it towards Malick and Giyera.

Restoring the Host's Body
"These are the healthiest we could find. Lucio has frozen them in place for you. I hope they're acceptable." "Yes. Well done." "One will be your host?" "I have a better use for them."

- Giyera and It

Using his memories, It looked back on the life of Grant Ward, noting that he wished they could have met under different circumstances, while Gideon Malick told It what a great soldier Ward was and how he had dedicated his life to HYDRA. It thanked Malick for arranging for Ward to travel to Maveth to die and be used as It's host, as Ward's body was the perfect host due to being strong and resilient.

Malick promised It that he was making moves to rebuild his army of Inhumans, explaining that he had recently found one for him. Lucio was brought to It with armed guards led by Giyera and Malick left them alone with the guards. It ordered Lucio's blindfold to be removed so he could see his eyes, allowing Lucio to use his powers against Giyera and the guards, paralyzing them. Unlike the paralyzed people, It was immune to Lucio's powers and used its abilities to get Lucio on his side. With Lucio now loyal to It, the Inhuman spent its time reading books on the history of humans until Malick arrived and asked to speak to It. Malick noted how Ward's corpse was not healing quickly enough and suggested that it was perhaps time for It to find a new host, although he made it clear he was not offering to give up his own body, suggesting instead Giyera or Lucio as a host. It made it clear that the host must be dead and he could not inhabit other Inhumans. Despite Malick offering to find a volunteer host, It refused and claimed Ward's body would serve his needs while Malick promised that there would be no more set-backs for their plans. When Malick left the base to attend a small summit in Taiwan, Hive listened in as Malick ordered Giyera to keep a close watch over Hive, as he was concerned over its condition. Giyera promised to do as instructed; however, Hive gave him new orders. Knowing that their true loyalty was now to him rather than to HYDRA Hive ordered Giyera and Lucio to leave his side and gather five humans for his own purposes, demanding that they be kept alive. Lucio paralyzed the five humans while Giyera confirmed that they were the healthiest that could be found, with It confirming that he approved. It told Giyera and Lucio that the people would not be his hosts as he had a better plan for them. Lucio commented that all these people were innocent, and It made it clear that he knew this was true, but they would now serve a greater purpose. It had Giyera and Lucio wait outside the room as It caused a sand tornado, disintegrating the kidnapped people's flesh and bodily fluids, causing the people to scream in agony. It used them to restore Grant Ward's body, healing his wounds, upon having reduced the five humans to skeletons while also being covered in a slimy substance.

Powers
"What are you?" "What are we?"

- Lucio and It

It is an Inhuman that achieved its genetic potential after undergoing Terrigenesis, granting It incredible superhuman powers. The transformation was physical, arguably the most drastic physical change ever seen in an Inhuman as It transitioned from being a human to a parasitic, worm-like organism.
 * Possession: It's Terrigenesis transformed it into a parasitic life form capable of infesting the bodies of the dead and using them as vessels for which it can perform basic actions. It possessed the corpse of an astronaut to try and fool Jemma Simmons, then possessed Daniels' corpse to try and trick Leo Fitz into letting it leave the planet. It can apparently also halt or slow the natural decomposition of a host body, given that it had been inhabiting Daniels' body for months. It also appears to have access to all of the hosts' memories and behavior, as it was able to successfully imitate Daniels during its earlier encounter with Fitz and relay information about Daniels' time with Simmons and also remembering the memories of love and anguish that his second host Grant Ward suffered.
 * Longevity: It's ability to inhabit the bodies of others has enabled it to survive for centuries.
 * Power Immunity: It has demonstrated to be immune to the powers of other Inhumans. It was unaffected by Lucio's paralyzing gaze, despite he had just paralyzed Giyera and two other guards restraining him.
 * Enhanced Smell: It can smell blood from a long distance away.
 * Environmental Manipulation: It had the ability to create sandstorms to confuse and isolate potential victims. It also widened a canyon to prevent Jemma Simmons and Will Daniels from reaching a wormhole, possibly by causing an earthquake.
 * Parallel Processing: It can process multiple sources of information at once. When It was catching up on historic events, after regaining its strength, It watched multiple documentaries at the same time.

Allies

 * HYDRA
 * Gideon Malick
 * Giyera
 * Grant Ward † - Host
 * Lucio

Enemies

 * Jemma Simmons
 * Leo Fitz
 * Phil Coulson
 * Project Distant Star Return Astronauts
 * Will Daniels † - Victim and Former Host
 * Austin † - Victim
 * Brubaker † - Victim
 * Taylor † - Victim
 * Manzini † - Victim

Appearances
In chronological order:
 * Agents of S.H.I.E.L.D.
 * Season Three
 * 4,722 Hours
 * Many Heads, One Tale (mentioned)
 * Closure (mentioned)
 * Maveth
 * Bouncing Back
 * The Inside Man
 * Parting Shot
 * Watchdogs

Trivia

 * In the comics, Hive is a parasitic entity created by HYDRA, composed of countless parasites merged around a host to form a single individual. Due to his intelligence, strength and dedication to the cause of HYDRA, Hive was appointed as one of HYDRA's heads by Wolfgang von Strucker.